1. Division of Cell Sciences, Institute of Comparative Medicine, Faculty of Veterinary Medicine, University of Glasgow, Glasgow, United Kingdom
2. Centre for Reproductive Endocrinology and Medicine, Division of Applied Medicine, Institute of Medical Sciences, University of Aberdeen, Aberdeen, United Kingdom
3. Macaulay Institute, Craigiebuckler, Aberdeen, UK
4. Unité de Biologie du Dévelopement et Reproduction, Institut National de la Recherche Agronomique (INRA), Jouy en Josas, France
5. Centre for Reproductive Biology, Queen’s Medical Research Institute, Edinburgh, United Kingdom