1. Department of Medicine and Bioregulatory Science, Graduate School of Medical Science, Kyushu University, Fukuoka, Japan
2. Department of Developmental Biology, National Institute for Basic Biology, Okazaki, Japan
3. Laboratory for Integrative Studies in Amphibian Biology, Group in Endocrinology, Museum of Vertebrate Zoology, Energy and Resources Group, and Department of Integrative Biology, University of California, Berkeley, California, USA
4. Graduate School of Medical Science, Kyushu University, Fukuoka, Japan