1. INBIO-UNELLEZ, Herbario PORT, Mesa de Cavacas, Guanare, estado Portuguesa, Venezuela; Instituto Venezolana de Investigaciones Científicas (IVIC), Caracas, Venezuela
2. Department of Botany, MRC-166, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, Washington DC 22013-7012, U.S.A.;