1. University of Florida Herbarium, Florida Museum of Natural History, Dickinson Hall, and Department of Biology, 379 1659 Museum Rd., Gainesville, Florida 32611, U.S.A.
2. Research Associate, Herbario Luis A. Fournier Origgi (USJ), Centro de Investigación en Biodiversidad y Ecología Tropical, Universidad de Costa Rica, Apdo. 11501-2060, San José, Costa Rica.
3. Department of Botany, California Academy of Sciences, San Francisco, California 94118, U.S.A.