1. Biodiversity Division, National Institute for Environmental Studies, Ibaraki 305–8506, Japan
2. Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Osaka 558–8585, Japan
3. Department of Natural History Science, Graduate School of Science, Hokkaido University, Sapporo 060–0810, Japan
4. Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa, 277–8564, Japan