1. Japan Fisheries Research and Education Agency, Fisheries Resources Institute, Fisheries Stock Assessment Center, Shimizu Field Station, 5–7–1 Orido, Shimizu-ku, Shizuoka, Shizuoka 424–8633, Japan
2. Laboratory of Environmental Zoology, Faculty of Agriculture, Meijo University, 1–501 Shiogamaguchi, Tenpaku-ku, Nagoya, Aichi 468–8502, Japan
3. Japan Fisheries Research and Education Agency, Fisheries Technology Institute, Nansei Branch, 422–1 Nakatsuhamaura, Minami-ise, Mie 516–0193, Japan