1. Division of Biodiversity, Institute for Agro-Environmental Sciences NARO, 3–1–3 Kannondai, Tsukuba-shi, Ibaraki 305–8604, Japan
2. 4–2–14 Matsubacho, Higashimatsuyama-shi, Saitama 355–0017, Japan
3. Division of Hilly and Semi-Mountainous Area Farming Research, Western Region Agricultural Research Center, NARO, 2575 Ikano-cho, Zentsuji-shi, Kagawa, 765–8508, Japan
4. Committee of Education of Hiroshima Prefecture, Naka-ku, Motomachi 9042, Hiroshima 730–8514, Japan