1. Department of Natural History Science, Graduate School of Science, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060–0810, Japan
2. Biodiversity Division, National Institute for Environmental Studies, Ibaraki 305–8506, Japan
3. Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa 277–8564, Japan
4. Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Osaka 558–8585, Japan