1. Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, U.K.
2. Department of Mathematics, Cavendish School of Computer Science, University of Westminster, 115 New Cavendish Street, London W1M 8JS, U.K.
3. Edward Grey Institute, Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, U.K.