1. Department of Natural Resources Conservation, 160 Holdsworth Way, University of Massachusetts, Amherst, MA 01003–9285, U.S.A.
2. Wildlife Conservation Society, 2300 Southern Boulevard, Bronx, NY 10460, U.S.A.
3. Institute of General and Experimental Biology, Mongolian Academy of Sciences, Ulaanbaatar 51, Mongolia
4. Ecology Department, National University of Mongolia, Ikh Surguuliin gudamj, Building 2, Ulaanbaatar, Mongolia