1. Ecology Group, Institute of Natural Resources, Massey University, Private Bag 11222, Palmerston North, New Zealand
2. School of Biological Sciences, University of Sydney, NSW 2006, Australia
3. Institute of Veterinary, Animal and Biomedical Sciences, Massey University, Private Bag 11222, Palmerston North, New Zealand
4. Statistics Research and Consultancy Centre, Massey University, Private Bag 11222, Palmerston North, New Zealand