1. Department of Biology, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen, Denmark
2. Land Resources and Environmental Sciences, Montana State University, Bozeman, Montana 59717
3. UN Environment World Conservation Monitoring Centre (UNEP WCMC), 219 Huntingdon Road, Cambridge, CB3 0DL, United Kingdom and Natural History, Museum of Denmark, Centre for Macroecology, Evolution & Climate, University of Copenhagen, Denmark