1. Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel email:
2. Institute for Biological Problems of the North RAS, Portovaya Str.18, Magadan, Russia and Zoological Museum, Biodiversity Unit, University of Turku, FI-20014, Finland and Department of Zoology & Entomology, University of the Free State, Bloemfontein