1. Department of Biochemistry, College of Medicine, The Catholic University of Korea, Seoul 06591, Korea
2. Institute for Aging and Metabolic Diseases, College of Medicine, The Catholic University of Korea, Seoul 06591, Korea
3. Department of Biomedicine & Health Sciences, College of Medicine, The Catholic University of Korea, Seoul 06591, Korea
4. Department of Mechanical Engineering, Johns Hopkins University, Baltimore, MD 21218, USA