1. Civil and Environmental Engineering, Imperial College London, London, UK
2. National Institute for Medical Research, Mwanza Center, Isamilo Street, Ilemela, Mwanza, Tanzania
3. College of Natural and Computational Sciences, Addis Ababa University, Arat Kilo, Addis Ababa, Ethiopia
4. Wolfson Wellcome Biomedical Laboratories, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK