1. Centre of International Research for Water and Environment (CIRSEE), Suez Lyonnaise des Eaux, 38, rue du Président Wilson, 78230, Le Pecq, France
2. School of Civil Engineering, Purdue University, West Lafayette, IN 47907-1284, USA
3. IFREMER Laboratoire de Microbiologie, DEL, BP 70, 29280 Plouzané, France