1. Institute of Wetland Research, Chinese Academy of Forestry, Haidian District, Beijing 100091, China; Beijing Key Laboratory of Wetland Services and Restoration, Institute of Wetland Research, Chinese Academy of Forestry, Haidian District, Beijing 100091, China; and Beijing Hanshiqiao National Wetland Ecosystem Research Station, Beijing 101309, China E-mail: lkyclj@126.com