1. a Aquatic Systems Research Group, School of Biology and Environmental Sciences, University of Mpumalanga, Nelspruit 1200, South Africa
2. b Stellenbosch Institute for Advanced Study (STIAS), Wallenberg Research Centre at Stellenbosch University, Stellenbosch 7600, South Africa
3. c Department of Geography and Environmental Sciences, University of Venda, Thohoyandou 0950, South Africa
4. d Institute for Global Food Security and School of Biological Sciences, Queen's University Belfast, Belfast BT9 5DL, UK