1. Kluyverlaboratory for Biotechnology, Delft University of Technology, Julianalaan 67, NL-2628 BC Delft, The Netherlands
2. Beijing University of Civil Engineering, 1 Zhalan Road, Xicheng district, Beijing 100044, China
3. Department of Microbiology, University of Nijmegen, Toernooiveld 1, 6525 ED Nijmegen, The Netherlands
4. Paques bv, P.O. Box 52, NL-8560 AB Balk, The Netherlands