1. a Research and Innovation Center, Institute of Technology of Cambodia, Russian Federation Blvd., P.O Box 86, Phnom Penh, Cambodia
2. b ISEM, Univ Montpellier, CNRS, IRD, CIRAD, Montpellier, France
3. c Bacteriology and Antibiotic resistance group, Medical Biology Laboratory, Institut Pasteur du Cambodge, Phnom Penh, Cambodia
4. d ISEM, Univ Montpellier, CNRS, IRD, Montpellier, France