1. Graduate School of Life Science, Hokkaido University, Kita-9, Nishi-9, Kita-ku, Sapporo 060-8589, Japan
2. Graduate School of Agriculture, Hokkaido University, Kita-9, Nishi-9, Kita-ku, Sapporo 060-8589, Japan
3. Graduate School of Environmental and Life Science, Okayama University, 2-1-1 Tsushima-Naka, Kita-ku, Okayama-shi, Okayama 700-8530, Japan
4. College of Agriculture, Academic Institute, Shizuoka University, 836 Ohya, Suruga-ku, Shizuoka-shi, Shizuoka 422-8529, Japan
5. Research Faculty of Agriculture, Hokkaido University, Kita-9, Nishi-9, Kita-ku, Sapporo 060-8589, Japan