1. Laboratory of Animal Hygiene, Joint Faculty of Veterinary Medicine, Kagoshima University, Kagoshima, Kagoshima, Japan
2. These authors contributed equally to this work
3. United Graduate School of Veterinary Science, Yamaguchi University, Yamaguchi, Yamaguchi, Japan
4. Transboundary Animal Diseases Center, Joint Faculty of Veterinary Medicine, Kagoshima University, Kagoshima, Kagoshima, Japan
5. Kagoshima Crane Conservation Committee, Izumi, Kagoshima, Japan