1. Dutch Wildlife Health Centre (DWHC), Utrecht University, Utrecht, The Netherlands
2. These authors contributed equally to the work
3. Veterinary Pathology Diagnostic Centre (VPDC), Division of Pathology, Department of Pathobiology, Utrecht University, Utrecht, The Netherlands
4. Sovon, Dutch Centre for Field Ornithology, Nijmegen, The Netherlands
5. Department of Animal Ecology, Institute for Water and Wetland Research, Radboud University Nijmegen, The Netherlands
6. Centre for Monitoring of Vectors (CMV), National Reference Centre (NRC), Netherlands Food and Consumer Product Safety Authority (NVWA), Ministry of Economic Affairs, Wageningen, The Netherlands
7. ErasmusMC, Department of Viroscience, Rotterdam, The Netherlands
8. Vogeltrekstation – Dutch Centre for Avian Migration and Demography (NIOO-KNAW), Wageningen, The Netherlands