1. Department of Population Health Sciences, School of Public Health, Georgia State University, Atlanta, Georgia, United States
2. Hakubi Center for Advanced Research, Kyoto University, Yoshidahonmachi, Sakyo-ku, Kyoto, Japan
3. Graduate School of Advanced Integrated Studies in Human Survivability, Kyoto University Yoshida-Nakaadachi-cho, Sakyo-ku, Kyoto, Japan
4. Seto Marine Biological Laboratory, Field Science, Education and Reseach Center, Kyoto University, Shirahama-cho, Nishimuro-gun, Wakayama, Japan
5. Department of Zoology, University of Oxford, Oxford, United Kingdom