1. These authors contributed equally to this work
2. Department of Virology, Faculty of Veterinary Medicine, Kaferelsheik University, Egypt
3. Department of Microbiology and Parasitology, College of Veterinary Medicine, King Faisal University, Saudi Arabia
4. School of Public Health, The University of Hong Kong, Hong Kong SAR, China
5. Centre of Influenza Research, The University of Hong Kong, Hong Kong SAR, China
6. Key Laboratory of Protein and Peptide Pharmaceuticals, Chinese Academy of Sciences - University of Tokyo Joint Laboratory of Structural Virology and Immunology, Beijing, China
7. HKU-Pasteur Pole, The University of Hong Kong, Hong Kong SAR, China
8. Food Animal Health Research Program, Veterinary Preventive Medicine Department, OARDC/The Ohio State University, Wooster, United States of America
9. Department of Clinical Studies, College of Veterinary Medicine, King Faisal University, Saudi Arabia