1. Department of Biological and Nutritional Sciences, University of Newcastle upon Tyne, Newcastle upon Tyne NE1 7RU, U.K.
2. Department of Physiological Sciences, University of Newcastle upon Tyne, Medical School, Newcastle upon Tyne NE2 4HH, U.K.
3. Department of Cellular Physiology, The Babraham Institute, Babraham, Cambridge CB2 4AT, U.K.