1. Blavatnik Institute of Cell Biology, Harvard Medical School, Boston, MA 02115, U.S.A.
2. The Broad Institute of MIT and Harvard, Cambridge, Massachusetts 02142, U.S.A.
3. Molecular Horizons and The School of Chemistry and Molecular Bioscience, University of Wollongong, Northfields Avenue, Wollongong, NSW 2522, Australia
4. Illawarra Health and Medical Research Institute, Northfields Avenue, Wollongong, NSW 2522, Australia