1. 1The Sainsbury Laboratory, University of East Anglia, Norwich Research Park, Norwich NR4 7UH, United Kingdom
2. 2Institute of Plant and Microbial Biology, Zurich-Basel Plant Science Center, University of Zurich, 8008 Zurich, Switzerland
3. 3RIKEN Center for Sustainable Resource Science, Yokohama, Kangawa 230-0045, Japan