1. Department of Medicine, University of Colorado Health Sciences Center, Denver, CO 80262, U.S.A.
2. Department of Immunology, University of Colorado Health Sciences Center, Denver, CO 80262, U.S.A.
3. Department of Biochemistry and Molecular Genetics, University of Colorado Health Sciences Center, Denver, CO 80262, U.S.A.
4. Department of Pharmacology, University of Colorado Health Sciences Center, Denver, CO 80262, U.S.A.
5. Department of Biochemistry and Molecular Biology, Royal Free and University College Medical School, University College London, London NW3 2PF, U.K.