1. Department of Biological Science, Graduate School of Science, Hiroshima University, Higashi-Hiroshima, 739-8526, Japan
2. Department of Stem Cell Biology and Histology, Graduate School of Medicine, Tohoku University, Sendai, 980-8575, Japan
3. Graduate School of Frontier Bioscience, Osaka University, Suita, 565-0871, Japan
4. Division of Chemistry, Faculty of Science, Hokkaido University, Sapporo, 060-0810, Japan