1. Laboratory of Molecular Biochemistry, School of Life Science, Tokyo University of Pharmacy and Life Science, 1432-1 Horinouchi, Hachioji, Tokyo 192-0392, Japan
2. Department of Biochemistry, Faculty of Medicine, University of Peradeniya, Peradeniya and Institute of Fundamental Studies, Kandy, Sri Lanka
3. Central Laboratories for Key Technology, Kirin Brewery Co., Yokohama, Kanagawa 236-0004, Japan
4. Department of Biology, Nippon Dental University, Chiyoda-ku, Tokyo 102-8159, Japan