1. Cranfield Health, Cranfield University, Cranfield, Bedfordshire MK43 0AL, U.K.
2. Laboratory of Chromatin and Gene Expression, The Babraham Institute, Babraham Research Campus, Cambridge CB22 3AT, U.K.
3. The Sir William Dunn School of Pathology, University of Oxford, South Parks Road, Oxford OX1 3RE, U.K.