1. Radboud University Nijmegen, ICIS, Heyendaalseweg 135, 6525 AJ, Nijmegen, The Netherlands
2. DOE Joint Genome Institute, Walnut Creek, CA 94598, U.S.A.
3. Department of Microbiology, Institute for Water and Wetland Research, Radboud University Nijmegen, Heyendaalseweg 135, 6525 AJ, Nijmegen, The Netherlands
4. Delft University of Technology, Department Biotechnology, 2628 BC, Delft, The Netherlands