1. Department of Biochemistry and Biophysics, University of Pennsylvania, Philadelphia, PA 19104, U.S.A.
2. Department of Biology, University of Pennsylvania, Philadelphia, PA 19104, U.S.A.
3. Penn Center for Genome Integrity, University of Pennsylvania, Philadelphia, PA 19104, U.S.A.
4. Epigenetics Institute, University of Pennsylvania, Philadelphia, PA 19104, U.S.A.