1. Department of Ophthalmology, Flinders University, Flinders Medical Centre, Adelaide, South Australia, Australia
2. Department of Ophthalmology, Flinders University, Flinders Medical Centre, Adelaide, South Australia, Australia2now also with Menzies Research Institute Tasmania, University of Tasmania, Tasmania, Australia
3. now also with Menzies Research Institute Tasmania, University of Tasmania, Tasmania, Australia3Centre for Eye Research Australia, Royal Victorian Eye and Ear Hospital, University of Melbourne, Melbourne, Victoria, Australia