1. Department of Pharmacology and Clinical Neuroscience, Umeå University, Umeå, Sweden
2. Institute of Neuroscience and Physiology, Department of Psychiatry and Neurochemistry, The Sahlgrenska Academy at University of Gothenburg, Mölndal, Sweden
3. Institute of Neuroscience and Physiology, Department of Psychiatry and Neurochemistry, The Sahlgrenska Academy at University of Gothenburg, Mölndal, Sweden3University College London Institute of Neurology, Queen Square, London, England