1. From the Wellcome Trust Centre for Cell-Matrix Research, Faculty of Life Sciences,
2. the School of Biomedicine, and the 4Manchester Academic Health Sciences Centre, Royal Eye Hospital, Central Manchester University Hospitals NHS Foundation Trust, Manchester, United Kingdom;
3. From the Wellcome Trust Centre for Cell-Matrix Research, Faculty of Life Sciences, the 5Department of Biochemistry, University of Oxford, Oxford, United Kingdom; and
4. the Materials Science Centre, University of Manchester, Manchester, United Kingdom;
5. the Department of Biochemistry, Radboud University Nijmegen Medical Center, Nijmegen, The Netherlands.