1. Department of Surgical Sciences, School of Veterinary Medicine, University of Wisconsin-Madison, WI, USA
2. Department of Ophthalmology and Visual Sciences, School of Medicine and Public Health, University of Wisconsin-Madison, WI, USA
3. Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin-Madison, WI, USA
4. Doheny Eye Institute, and Department of Ophthalmology University of California, Los Angeles, CA, USA