1. Department of Ophthalmology and Visual Sciences, The University of Iowa, Iowa City, IA, USA
2. Center for the Prevention and Treatment of Visual Loss, Iowa City VA Health Care System, Iowa City, IA, USA
3. Department of Biomedical Research, National Jewish Health, Denver, CO, USA
4. Department of Immunology & Microbiology, University of Colorado Anschutz Medical Campus, Aurora, CO, USA
5. Iowa Institute of Human Genetics, Carver College of Medicine, The University of Iowa, Iowa City, IA, USA
6. Roy J. Carver Department of Biomedical Engineering College, The University of Iowa, Iowa City, IA, USA