1. From the Jackson Laboratory, Bar Harbor, Maine;
2. the Cole Eye Institute and
3. the Department of Neuroscience, Lerner Research Institute, Cleveland Clinic Foundation, Cleveland, Ohio;
4. the Cole Eye Institute and the 4Research Service, Cleveland VA Medical Center, Cleveland, Ohio; and the 5Department of Ophthalmology, Cleveland Clinic Lerner College of Medicine, Case Western Reserve University, Cleveland, Ohio.