1. From the Department of Ophthalmology, University of Washington School of Medicine, Seattle, Washington;
2. Cole Eye Institute and Lerner Research Institute, Cleveland Clinic Foundation, Cleveland, Ohio;
3. Department of Molecular Biology and Genetics, Cornell University, Ithaca, New York; and the
4. From the Department of Ophthalmology, University of Washington School of Medicine, Seattle, Washington;4Department of Biochemistry, University of Washington, Seattle, Washington.