1. Experimental Therapeutics Centre, Agency for Science, Technology and Research (A*STAR), Singapore
2. Singapore Eye Research Institute, Singapore
3. Institute of Medical Biology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore
4. Experimental Therapeutics Centre, Agency for Science, Technology and Research (A*STAR), Singapore 4Department of Physiology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore