1. Department of Ophthalmology, Flinders University, Adelaide, South Australia, Australia
2. Richard Lindsay and Associates, East Melbourne, Victoria, Australia
3. Centre for Vision Research, Westmead Millennium Institute and University of Sydney, Sydney, New South Wales, Australia
4. Centre for Vision Research, Westmead Millennium Institute and University of Sydney, Sydney, New South Wales, Australia 4Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, Melbourne, Victoria, Australia