1. Department of Ophthalmology, Wakayama Medical University School of Medicine, Wakayama, Japan
2. Program in Genes and Development, The University of Texas Graduate School of Biomedical Sciences at Houston, Houston, Texas, United States 3Department of Genetics, University of Texas MD Anderson Cancer Center, Houston, Texas, United States
3. The Laboratory Animal Center, Wakayama Medical University School of Medicine, Wakayama, Japan