1. University College London Institute of Ophthalmology, London, United Kingdom
2. Clinical Ophthalmology at the Centre for Vision and Vascular Science, Queens University Belfast, Belfast, Northern Ireland
3. University College London Institute of Ophthalmology, London, United Kingdom 2Moorfields Eye Hospital, London, United Kingdom
4. Department of Ophthalmology, Institute of Medical Sciences, University of Aberdeen, Aberdeen, Scotland, United Kingdom
5. Laboratory of Visual Physiology, National Institute of Sensory Organs, National Hospital Organization, National Tokyo Medical Center, Meguro-ku, Tokyo, Japan
6. Department of Ophthalmology, Keio University School of Medicine, Shinjyuku-ku, Tokyo, Japan
7. University of Iowa, Institute for Vision Research, Howard Hughes Medical Institute, Iowa City, Iowa