1. Department of Public Health and Primary Care, Institute of Public Health, University of Cambridge School of Clinical Medicine, Cambridge, United Kingdom
2. Division of Genetics and Epidemiology, University College London Institute of Ophthalmology, London, United Kingdom
3. Department of Ophthalmology, Norfolk & Norwich University Hospital, Norwich, United Kingdom
4. National Institute for Health Research Biomedical Research Centre, Moorfields Eye Hospital National Health Service Foundation Trust and University College London Institute of Ophthalmology, London, United Kingdom
5. Division of Genetics and Epidemiology, University College London Institute of Ophthalmology, London, United Kingdom 4National Institute for Health Research Biomedical Research Centre, Moorfields Eye Hospital National Health Service Foundation Trust and Un