1. Centre for Eye Research Australia, Royal Victorian Eye and Ear Hospital, Melbourne, Victoria, Australia 2Department of Ophthalmology, Keio University, Tokyo, Japan
2. Centre for Eye Research Australia, Royal Victorian Eye and Ear Hospital, Melbourne, Victoria, Australia 3Department of Public Health, Yamagata University Faculty of Medicine, Yamagata, Japan
3. Centre for Eye Research Australia, Royal Victorian Eye and Ear Hospital, Melbourne, Victoria, Australia
4. Centre for Eye Research Australia, Royal Victorian Eye and Ear Hospital, Melbourne, Victoria, Australia 4Singapore Eye Research Institute, National University of Singapore, Singapore
5. Centre for Eye Research Australia, Royal Victorian Eye and Ear Hospital, Melbourne, Victoria, Australia 5Centre for Vision Research, Westmead Millennium Institute and Department of Ophthalmology, University of Sydney, Westmead Hospital, Sydney, New South