1. Department of Surgical and Radiological Sciences, School of Veterinary Medicine, University of California-Davis, Davis, CA, USA
2. Department of Ophthalmology and Visual Sciences, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan
3. Center for Health and the Environment, University of California-Davis, Davis, CA, USA
4. Department of Anatomy, Physiology and Cell Biology, School of Veterinary Medicine, University of California-Davis, Davis, CA, USA
5. Department of Ophthalmology & Vision Science, School of Medicine, University of California-Davis, Davis, CA, USA