1. UCL Institute of Ophthalmology, University College London, London, United Kingdom
2. Moorfields Eye Hospital, London, United Kingdom
3. Section of Academic Ophthalmology, School of Life Course Sciences, FoLSM, King's College London, United Kingdom
4. Department of Twin Research and Genetic Epidemiology, School of Life Course Sciences, FoLSM, King's College London, London, United Kingdom
5. Jones Eye Institute, Department of Ophthalmology, University of Arkansas for Medical Sciences, Little Rock, Arkansas, United States
6. Centre for Ophthalmology and Visual Sciences (Incorporating Lions Eye Institute), The University of Western Australia, Perth, Western Australia, Australia
7. Ophthalmology, Department of Surgery, University of Melbourne, Melbourne, Victoria, Australia
8. Department of Ophthalmology and Visual Sciences, Medical College of Wisconsin, Milwaukee, Wisconsin, United States
9. Department of Cell Biology, Neurobiology and Anatomy, Medical College of Wisconsin, Milwaukee, Wisconsin, United States