1. Department of Biochemistry and Molecular Biology, Sidney Kimmel Medical College at Thomas Jefferson University, Philadelphia, PA, USA
2. Department of Ophthalmology, Wills Eye Hospital, Sidney Kimmel Medical College of Thomas Jefferson University, Philadelphia, PA, USA
3. Department of Pathology, Wills Eye Hospital, Sidney Kimmel Medical College of Thomas Jefferson University, Philadelphia, PA, USA
4. PXE International Center of Excellence in Research and Clinical Care, Thomas Jefferson University, Philadelphia, PA, USA
5. Bower Laboratory for Translational Medicine Vickie and Jack Farber, Vision Research Center at Wills Eye Hospital, Wills Eye Hospital, Philadelphia, PA, USA